Snow White and Charming need to give up their hearts if they want to save Storybrooke from the Evil Queen. A magical sapling is sought. Snow White tries to get away from the Woodcutter. David has plans to sell the family farm.
The Evil Queen threatens to unleash water from the river of lost souls on everyone in Storybrooke, unless Mary Margaret and David hand over their shared heart. Regina tries to put Zelena against the Evil Queen, as flashbacks show Snow White and David's paths cross before she robs Abigail
In Storybrooke, the Evil Queen threatens to destroy the town and everyone in it unless Snow and David surrender their hearts. As Emma, Hook, and Henry prepare to defend Storybrooke, Snow, David, and Regina search for a magical sapling created by the first spark of true love.